They realise that if their universe was a 4-torus, that would
result in extra modes for fermionic vacuum energy that led to an
overall negative energy density (fermionic vacuum energy is negative)
which, in this kind of universe, would require space to be positively
curved everywhere. But you can't have a space with the topology of
a 4-torus that is positively curved everywhere. So what they conclude
is that the universe being a 4-torus is self-contradictory.
However ...
They have also known for a long time that the universe must be finite
in all directions, to avoid exponentially growing solutions to the
wave equation. So the simplest alternative topology to a 4-torus is
a 4-sphere. In that case, the topology doesn't have the extra
fermionic modes, and the overall vacuum energy is positive, which, in
this kind of universe, requires space to be negatively curved.
So ...
It's not that they conclude that the curvature actually is positive
everywhere, and hence the universe can't be a 4-torus. It's that they
see why a 4-torus both implies positive curvature and at the same time
is ruled out by positive curvature, which eliminates the whole possibility.
You ask:
Also, why can't the curvature be negative everywhere,
resulting in a hyperbolic universe?
This universe ...
can't be an infinite hyperbolic universe, which is what is
usually meant by that phrase. However, you can have a finite universe
with the topology of a 4-sphere but negative curvature.
It just can't be uniform negative curvature, it has to vary in
magnitude from place to place.
More details at http://www.gregegan.net/ORTHOGONAL/06/GRExtra.html